Showing posts with label Pleistocene. Show all posts
Showing posts with label Pleistocene. Show all posts

Monday, 20 June 2016

Where did the wild horse go?

Tarpan. Isn't it an evocative word? It brings to mind images of wild horses, untamed by man, of fierce stallions and galloping herds. When it comes to wild horses in Europe, "Tarpan" is the go to term. Unfortunately, it is also one shrouded in controversy, and for one simple reason: nobody really knows what it means. It seems simple enough - the Tarpan is another word for the European wild horse, right? Well... kinda. Sometimes. It is certainly true that the undomesticated horses which roamed Europe in prehistory are commonly referred to as Tarpans, but the problem is that the designation is not exclusively limited to them - nor necessarily should it be. If Tarpan is a word for the European wild horse, then it should apply to all members of that species, regardless of when they lived. The factor that complicates everything and throws this neat and tidy definition out of the window can be summarized neatly in one word: Interbreeding.

Europe has been home to domestic horses for a long time, and for the entirety of their time coexisting with their wild relatives, interbreeding has most likely occurred. Now, at first this was not an issue, since the populations of (mostly) pure wild horses were so larger that the genes of a few domestics did not have much of an impact. As time went on however, and the wild horse populations plummeted, this genetic balance was lost. Suddenly there were significantly more domestic horses than wild ones, and in times of war or simply due to accidents, domestics were often abandoned and left to their own devices, resulting in them becoming feral. This is where the boundary between wild horses and domestics becomes incredibly blurred, and where the word Tarpan really starts to grow sketchy. Historical observers from centuries passed were rarely detail-orientated enough to take note of the subtle morphological distinctions that separated wild horses and domestics, so whenever a herd of wild-living horses was spotted, these animals were simply referred to as wild, and sometimes, you guessed it, "Tarpans". This makes it very difficult to ascertain whether any given historical account of wild horses living in an area actually refer to true, undomesticated horses, or simply feral descendants of domestics. To make matters even more complicated, as mentioned before, extensive interbreeding took place, and in the latter years of the wild horse's existence, it is plausible if not almost certain that very nearly all populations of so-called Tarpans were either hybrids or pure domestic animals, with very few pure wild types remaining. This is often referred to as the only known picture of a Tarpan, but looking at its morphology, the animal shown is almost certainly either a hybrid or a pure domestic, as it bares very little resemblance to neither the skeletons we have of European wild horses, nor the extant and closely related Przewalski's horses.

To add one final layer of complexity and doubt to the word "Tarpan", today it is often used to refer to certain "primitive" breeds of domestic horses, particularly the Konik Polski. Koniks are one of several breeds often claimed to be either direct descendants of wild horses, or, in the Konik's case, a deliberate back-breeding attempt that incorporated hybrids of wild and domestic animals, thus resulting in a breed almost identical to the original wild horse. While the Konik is undeniably a sturdy and "wild"-looking breed, its morphology, like that of the animal in the picture claimed to be a Tarpan, and all other modern horse breeds, does not match that of the wild horse. Genetic tests have shown that it is no more closely related to wild horses than any other modern breed, and the story of it being a back-breeding attempt has been demonstrated to be a myth. In reality, the Konik is simply a particularly hardy breed originally used as draft animals in their native Poland, where the breed presumably originated, which just so happens to somewhat resemble the original wild horse. And yet, regardless of everything I just told you, the Konik is still often referred to as a Tarpan, including by organizations such as Rewilding Europe.

Pictured: Not Tarpans, but instead a herd of Przewalski's horses, the world's only still extant species of wild horse.


Friday, 27 November 2015

Elephants Of Antiquity

Elephants, members of the group proboscidea, are beloved by everyone; they are big, charismatic, and essential keystone species throughout their native range, even in the places where they are now extinct. Once one of the most widely distributed groups of large herbivorous mammals on the planet, they are now confined to tropical Asia and Africa, and even then just barely in many places. I have mentioned them before when talking about some of the regions which they once inhabited, but never in great detail. That changes now, and in this post I hope to cover pretty much the entire native range of proboscideans on all continents, focusing mostly on those now extinct. Just to clarify briefly what I mean when I say "native", while the term has no true agreed upon meaning, for the sake of this post I will define it as any animal which is currently or would still have been present in an area, if not for human activity. Another fact that must be mentioned is that the terms "proboscidean" and "elephant" do not mean the same. All elephants are proboscideans, but not all proboscideans are elephants. The proboscidean family tree is typically divided into two main groups, the elephants and the mastodons, which split early on in their history. So, with that settled, let us begin with the two places where elephants are still present.

Proboscidean cladogram, by Vladimir Nikolov