Where did the wild horse go?

Tarpan. Isn't it an evocative word? It brings to mind images of wild horses, untamed by man, of fierce stallions and galloping herds. When it comes to wild horses in Europe, "Tarpan" is the go to term. Unfortunately, it is also one shrouded in controversy, and for one simple reason: nobody really knows what it means. It seems simple enough - the Tarpan is another word for the European wild horse, right? Well... kinda. Sometimes. It is certainly true that the undomesticated horses which roamed Europe in prehistory are commonly referred to as Tarpans, but the problem is that the designation is not exclusively limited to them - nor necessarily should it be. If Tarpan is a word for the European wild horse, then it should apply to all members of that species, regardless of when they lived. The factor that complicates everything and throws this neat and tidy definition out of the window can be summarized neatly in one word: Interbreeding.

Europe has been home to domestic horses for a long time, and for the entirety of their time coexisting with their wild relatives, interbreeding has most likely occurred. Now, at first this was not an issue, since the populations of (mostly) pure wild horses were so larger that the genes of a few domestics did not have much of an impact. As time went on however, and the wild horse populations plummeted, this genetic balance was lost. Suddenly there were significantly more domestic horses than wild ones, and in times of war or simply due to accidents, domestics were often abandoned and left to their own devices, resulting in them becoming feral. This is where the boundary between wild horses and domestics becomes incredibly blurred, and where the word Tarpan really starts to grow sketchy. Historical observers from centuries passed were rarely detail-orientated enough to take note of the subtle morphological distinctions that separated wild horses and domestics, so whenever a herd of wild-living horses was spotted, these animals were simply referred to as wild, and sometimes, you guessed it, "Tarpans". This makes it very difficult to ascertain whether any given historical account of wild horses living in an area actually refer to true, undomesticated horses, or simply feral descendants of domestics. To make matters even more complicated, as mentioned before, extensive interbreeding took place, and in the latter years of the wild horse's existence, it is plausible if not almost certain that very nearly all populations of so-called Tarpans were either hybrids or pure domestic animals, with very few pure wild types remaining. This is often referred to as the only known picture of a Tarpan, but looking at its morphology, the animal shown is almost certainly either a hybrid or a pure domestic, as it bares very little resemblance to neither the skeletons we have of European wild horses, nor the extant and closely related Przewalski's horses.

To add one final layer of complexity and doubt to the word "Tarpan", today it is often used to refer to certain "primitive" breeds of domestic horses, particularly the Konik Polski. Koniks are one of several breeds often claimed to be either direct descendants of wild horses, or, in the Konik's case, a deliberate back-breeding attempt that incorporated hybrids of wild and domestic animals, thus resulting in a breed almost identical to the original wild horse. While the Konik is undeniably a sturdy and "wild"-looking breed, its morphology, like that of the animal in the picture claimed to be a Tarpan, and all other modern horse breeds, does not match that of the wild horse. Genetic tests have shown that it is no more closely related to wild horses than any other modern breed, and the story of it being a back-breeding attempt has been demonstrated to be a myth. In reality, the Konik is simply a particularly hardy breed originally used as draft animals in their native Poland, where the breed presumably originated, which just so happens to somewhat resemble the original wild horse. And yet, regardless of everything I just told you, the Konik is still often referred to as a Tarpan, including by organizations such as Rewilding Europe.

Pictured: Not Tarpans, but instead a herd of Przewalski's horses, the world's only still extant species of wild horse.

A thought experiment in the Highlands

Much ink has been spilled over the merits and flaws of rewilding. It is a concept which I have come out in support of multiple times before, though I take issue with many of the claims and arguments made by some so-called "rewilders". One of the main points of disagreement between advocates and opponents of rewilding has been as to what impacts simply letting nature reassert itself would have. Some people say that it would be negative, that human activities are necessary for the preservation of our ecosystems, while others say that it is unquestionably a positive, that can only serve to better our wildlife. Both of these positions are of course extremes, and most people lie somewhere in between, but they give a  general idea of what the sides are. To actually answer the question is a difficult if not impossible task, as it would depend on a wide variety of factors. Whether reintroductions are considered a necessary part of rewilding, for instance, completely changes what results said process would have. In this post, I will indulge in a thought experiment of sorts, centered on the British highlands: What if all farming along with most other human activity everywhere in the uplands was stopped, and deer numbers were heavily culled and kept at bay. No reintroductions, no direct action aside from the culling, just a complete ceasing of all activities in the highlands. I do not actually advocate this scenario, nor do I find it remotely plausible, but it offers some idea of what the outcome of one extreme form of rewilding would be. As we will eventually see, it also shows why reintroductions are in fact necessary for rewilding and habitat restoration in the long term, and that "grazing bad, trees good" is not always true.

The British highlands as they appear today, beautiful but lifeless

Exploring the natural land cover of Europe

How would Europe have looked if we had never come along? This subject has been covered before, specifically as goes the fauna, but today I will be tackling a different side of this, namely how the landscape itself would have looked. Traditionally imagined as a dense, continent-wide closed canopy forest, this view now seems unlikely, but precisely how it looked is difficult to assess, simply due to our lack of clear evidence. Despite this, enough data is available to give us a general picture of roughly how it may have looked, even if a large amount of educated guesses and inferences are required. The importance of discussing this subject must not be understated, as it is absolutely vital to conservation and habitat restoration. As has been covered here, trying to protect and restore the wrong habitats can have a devastating effect on wildlife, as demonstrated by the stark decline in many species caused by the intensive management of heaths throughout much of the continent. To understand how best to protect species we must first know what their natural habitats were, and in what conditions they once thrived. In this post, I will be covering the continent in general, but again with a stronger focus on western Europe, as this is where most attempted habitat restoration is currently taking place and where data is most plentiful.

Old-growth beech forest in Slovakia, often but mistakenly used as a model for all pristine European habitats

Lands Pristine: Europe during the Eemian

Previously I have spoken about prehistoric Australia in great length, namely here and here, and recently wrote an in-depth post about ancient California. Pre-Human Europe however has been something of a neglected subject, despite also being the one I have the greatest interest in. I have mentioned details about it in passing, but now is its time in the limelight.

When people think of prehistoric Europe, they typically picture the early Holocene, covered here, but as that post discusses, the pristine European wilderness was already gone by then. No, to see Europe as it would still have looked today if not for our presence, one must go back to the Eemian, also known as the last interglacial. Spanning from 130,000 to 115,000 years ago, it was the last point in time when the climate resembled todays, but the megafauna had not yet died out. All of the modern species we know today already existed, and at a cursory glance it would have looked quite similar, but the Eemian was to modern Europe what present day southern Africa is to its northern counterpart, an unspoilt, pristine version of the latter, in which the now vanished giants were not just present, but dominated the landscape. In this post I will be covering primarily western Europe, as that is the place with which I am most familiar, and we will explore a Europe quite different from what we know today.

Trafalgar Square, 130,000 years ago, by Roman Uchytel

Big cats of Europe - past and present

British big cats, German big cats, Danish big cats, French big cats. Every European country seems to be reporting sightings of mysterious, large, often black cats nowadays. Of course, in the vast majority of cases, these are either hoaxes or misidentification, the fact that melanistic animals are most often reported, despite being far rarer than individuals of normal colouration, should itself be a clue to this reality. Whatever the case. some of these sightings have possibly been of real big cats. In central Europe for instance, lynx are fairly widespread, and to a person unfamiliar with the animal, could quite convincingly appear as a leopard or jaguar. These sightings are less tenable on the British isles, but even in Scotland wildcats are present, if barely, and hybrids between them and domestics are often significantly larger than your average housecat. Barring the occasional zoo or circus escapee however, the only true big cats to be encountered in present day Europe is the aforementioned lynx. But this was not always the case. Today I will be talking about the history of big cats in Europe since the last interglacial, their distribution, diversity, success, and eventual downfall by way of spears and bows.

An Eurasian lynx, the only real big cat left in Europe

The early Holocene of Europe - from 11,000 to 6000 BP

The Quaternary extinction is a much talked about subject today, as it is both fascinating in the context of prehistory, and simultaneously important in modern conservation. Typically defined as the rapid decline in global biodiversity during the late Pleistocene up until present, it is sometimes referred to as the sixth mass extinction in earth's history, the previous one wiping out the non-avian dinosaurs 66 million years ago. Personally, I dislike the term Quaternary extinction. The event mostly occurred within the last 50,000 years, making the term "Quaternary" odd, since the Pliocene and majority of the Pleistocene was not involved. Some have also referred to it as the Pleistocene extinction, which is equally incorrect, as it has continued up until the present. I prefer the term "Anthropogenic extinction", and have used it previously, as I believe it is more apt. It ignores the timeframe in which it has occurred, instead describing the primary causal factor, humans.

While it is true that the most dramatic of the Anthropogenic extinctions occurred either during the late Pleistocene or around the industrial revolution, in this post I will be focusing on a very different, but in some ways even more significant period, the early Holocene. The early Holocene is special in that it represents a relatively brief span of time, approximately 5000 years, in which environmental conditions were similar to today, yet direct human influence on the landscape was not. Instead, the most major human influences during the early Holocene were the same as those during the Pleistocene - hunting. While many of the most important megafauna were already gone by the time the Holocene began, several were not, and for just a few thousand years, we got a glimpse of something similar to what the world may have looked like had humans not existed.

The world that we forgot

Humans are an inherently short sighted species. We do not think too far into the future, and rarely gaze much further back than our youth. At most, the average person thinks a few decades forward, and a century back, anything beyond that is largely forgotten, or ignored. For conservationists protecting nature, this cone of vision is slightly less narrow, often looking an entire century forward, and sometimes upwards of 500 years back. This allows them to view the general trends these last few centuries have experienced. But it is still a narrow view, and still does not offer one crucial thing; Perspective. Most conservation efforts focus either on preserving areas as they are today, or turning them back at most a few hundred years. Often the time just before the industrial revolution is used as a target, an ideal. In this post I will discuss why to properly conserve nature, one must look much further back, and how this lack of perspective and vision is choking nature today.

The Cairngorms of eastern Scotland